Tully Jr (Anson Mount) and his younger brother Earl (Glenn Fitzgerald) live with their distant father on their ranch in Nebraska. While the shy, sensitive Earl spends his free time watching movies, the more gregarious Tully works his way through a succession of short-term affairs and maintains an on-off relationship with April (Catherine Kellner), a stripper in town. However, when their childhood friend Ella (Julianne Nicholson) returns to town to start a veterinary practice, Tully finds himself falling in love with her - although the townspeople are doubtful that he could ever commit to one woman. Meanwhile, their father, Tully Sr, struggles with a brooding sadness, a clue to past wounds taking too long to heal. When several secrets surface, the family dynamic is changed forever.